The player then assumes the role of Flight Officer Paul VanAtta from the Eighth Air Force, who bombs Nijmegen while defending the B-24 Liberators from German fighters. After the glider crashes near Carentan the squad regroups with other glider teams and attack a nearby town. The player then plays as Private Jonathan Shepherd. The player then goes to France with Daniel Ferguson, where they battle the Germans at Ste. The game begins in Italy where the player, playing as Private Derrick Warren from the 82nd Airborne Division, takes part in an attack to recapture Altavilla from the Germans. In the British Campaign, the player fights in the Parachute Regiment during 'Operation Market Garden' and 'Operation Varsity'. As for the Canadians Campaign, the player takes the role of various soldiers of the 1st Canadian Army during the 'Battle of the Scheldt', 'Operation Infatuate', and 'Operation Blockbuster'.

The American Campaign takes place in the perspective of a soldier from the 82nd Airborne Division in 'Operation Avalanche', 'Operation Detroit', and 'Operation Market Garden'. Call of Duty: Roads to Victory lets the player play in three campaigns: American, British, and Canadian.

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